Whole-Wheat Bread With Walnuts, Sesame, And Pumpkin Seeds

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1 package dry yeast (2 1/4 teaspoons)

1 teaspoon sugar

1/2 cup warm water (100° to 110°)

1 1/2 cups plain low-fat yogurt

6 tablespoons honey

1/4 cup butter, melted and cooled

2 1/2 cups whole-wheat flour

3/4 cup chopped walnuts

1/2 cup plus 2 teaspoons pumpkin seeds, divided

1/2 cup plus 2 teaspoons sesame seeds, divided

1 tablespoon salt

2 1/4 cups all-purpose flour, divided

Cooking spray

1 large egg white, lightly beaten

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